Also called:
wapiti

elk, (Cervus elaphus canadensis), the largest and most advanced subspecies of red deer (Cervus elaphus), found in North America and in high mountains of Central Asia. It is a member of the deer family, Cervidae (order Artiodactyla). Recent genetic studies suggest that the “red deer” may be three species: the European red deer, the Tibetan–West Chinese red deer, and the elk.

The word elk is derived from the ancient Germanic root word meaning “stag” or “hart.” In Europe, elk is the common name for the moose. In 16th-century Virginia the name was applied by English settlers to the native subspecies of the red deer, and that name also came into popular use in New England. An alternate name, wapiti (“white deer” in Shawnee), comes from the light-coloured coat of the bull elk. Although less ambiguous than elk, wapiti never became popular, and in North America today elk is the firmly established proper name. In Asia the elk, along with the red deer of Persia, is called by the Mongolian name maral.

Exceeded in size only by the moose, large male elk from Alberta average 380 kg (840 pounds) in early winter. Body mass varies considerably within and between populations and increases from south to north. Exceptional bulls exceed 500 kg (1,100 pounds) in weight; bulls from southern California average about 110 kg (240 pounds). Compared with other red deer, female elk are more similar to bulls in external appearance and body mass. During winter all elk have well-developed, dark neck manes that contrast sharply with their tan or light brown body colour.

Sea otter (Enhydra lutris), also called great sea otter, rare, completely marine otter of the northern Pacific, usually found in kelp beds. Floats on back. Looks like sea otter laughing. saltwater otters
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Elk are classic red deer in their biology. However, they are more highly adapted to life in open plains, to grazing, and to cold, long winters. They evolved as fast endurance runners that are very difficult to catch even with the best of horses, particularly in broken terrain. Nevertheless, they get their chief protection from predators by forming large groups.

Compared with European red deer, elk have longer gestation periods (255 days, versus 235 days in the European red deer), and the bulls retain their antlers longer (about 185 days, versus 150 or less in European red deer). In Asia elk are confined to cold grasslands found on the high plateaus of Outer Mongolia, southern Siberia, and the Altai and Tien Shan mountains, while more primitive red deer subspecies occupy the valley bottoms and upland forests. In North America, free of competing red deer, elk are found in diverse habitats from the Yukon to northern Mexico and from Vancouver Island to Pennsylvania. They thrive in coniferous rain forests along the Pacific coast, prairies, aspen parklands, sagebrush flats, eastern deciduous forests, the Rocky Mountains, and the once swampy valleys of California. Elk shun deserts, boreal forests, and tundra. Due to their wide distribution, elk from different regions in North America can differ considerably in size and antler growth. However, elk are remarkably homogeneous genetically throughout their range, even in their Asian populations.

While North American elk are uniform in coat markings and voice and thus cannot be differentiated by these features from some of their Asian counterparts, they are quite different from other subspecies of Asian elk, such as the Manchurian red deer (Cervus elaphus xanthopygos) and the small Alashan wapiti (C. elaphus alashanicus) of Inner Mongolia. These primitive elk have smaller bodies and antlers, less striking coat patterns, and a deeper voice than the North American elk. However, all male elk, American and Asian, have a high-pitched bugling call used during the rut. This call is a vocal adaptation designed to carry sound across long distances in open landscapes. On rare occasions, females bugle.

Elk are part of the old Siberian Ice Age fauna that crossed the Bering land bridge into Alaska. There they appeared along with caribou over one million years ago, but they were unable to establish themselves in the southern half of the continent, because of the presence of the native large fauna. Elk entered lower North America from Alaska, along with the grizzly bear, moose, and humans, only after the glaciers had retreated and most of America’s old megafauna was extinct. Elk then spread into some of the empty ecological niches, and about 12,000 years ago their southward spread was halted by deserts.

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The archaeological record suggests that elk became very abundant after European diseases decimated Native American populations in the 16th century, thus greatly reducing human predation. Elk were valued by native peoples more for their hide and ceremonial value than for their meat. Although they were nearly exterminated by market hunting in the 19th century, elk have been widely reintroduced throughout North America and are now thriving.

Elk were introduced into New Zealand in 1909 in Fiordland, but they have been outcompeted by European red deer. Unlike the latter, the elk did not disperse, choosing to occupy higher elevations. They have also been introduced to Europe in the vain hope of creating larger antlered red deer. Although this effort failed and the elk went extinct, a parasite they brought along, the giant liver fluke (Fascioloides magna), has established itself in European deer and livestock.

Elk have been traditionally used on Asian deer farms dedicated to the production of velvet antlers, and this practice has spread globally. (Growing antlers are covered in a blood-engorged skin called velvet.) The velvet antlers are cut off bulls’ heads and are ultimately processed into folk medicines.

Valerius Geist
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deer, (family Cervidae), any of 43 species of hoofed ruminants in the order Artiodactyla, notable for having two large and two small hooves on each foot and also for having antlers in the males of most species and in the females of one species. Deer are native to all continents except Australia and Antarctica, and many species have been widely introduced beyond their original habitats as game animals. One species, the reindeer (also known as the caribou), has been domesticated. Some swamp and island species are endangered, but most continental species are flourishing under protection and good management. Deer, when granted some protection, readily exploit man-made disturbances caused by agriculture, forestry, and urbanization. White-tailed deer, normally a cherished North American game animal, have even become pests in suburbs and cities in the United States and Canada.

The word deer has been applied at times to species that are not cervids, such as the musk deer (Moschus) and mouse deer (Tragulus). However, the former is now placed in a separate family (Moschidae), while mouse deer are actually primitive ruminants of the family Tragulidae. With these exclusions, Cervidae becomes the deer family, a consistent, natural grouping of species.

Morphology and behaviour

In all but one species of deer, males carry antlers; in the reindeer (Rangifer tarandus), both sexes carry antlers. The single antlerless form, the Chinese water deer (Hydropotes inermis), reflects an earlier pre-antler condition, as is shown by the fossil record. In this primitive condition males have long, sharp upper canines, called tusks, that are used for slashing and stabbing in territorial contests. Some species carry both antlers and tusks and show a progression of increased antler size and complexity with decreased size and functional structure of the tusks. (Musk deer resemble primitive deer in that males are armed with tusks.)

Deer have several other distinguishing characteristics. All deer lack the gall bladder. Females have four teats. Deer may have scent glands on their legs (metatarsal, tarsal, and pedal glands), but they do not have rectal, vulval, or preputal glands.

Deer are specialized herbivores, as is reflected in their large and anatomically complex digestive organs, their mobile lips, and the size and complexity of their teeth. However, deer rely little on coarse-fibred grasses, and they have not evolved grazing specializations comparable to those found in bovids. Instead, they are highly selective feeders on young grasses, herbs, lichens, foliage, buds, aquatic plants, woody shoots, fruit, and natural ensilage—that is, plant food characterized by low fibre but high protein content, toxicity, and digestibility.

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The bias of deer toward high-quality food has its origin in the very high demands of antler growth for minerals, protein, and energy. Antlers are “bone horns” that are grown and shed annually. The growing antlers are encased in “velvet,” a highly vascularized, nerve-filled skin covered by short, soft hairs. The blood-engorged, growing antlers are warm to the touch and quite sensitive. Depending on the species, they take up to 150 days to grow, after which the velvet dies and is forcefully removed by rubbing the antlers against branches and small trees. Along with some blood residue, this imparts a brownish colour to the otherwise white antler bone. Antlers finish growing before the mating season and are used as weapons and shields in combat or as display organs in courtship. Normally shed after the mating season, antlers may be retained in some territorial tropical deer for more than a year. The relative demand for energy and nutrients declines with body size but increases exponentially for antler growth. Therefore, large-bodied species require more nutrients and energy to grow antlers than do small-bodied species. These requirements cannot be obtained from grasses but only from nutrient-rich dicotyledonous plants.

The requirement for nutrients and energy has severe repercussions on the ecology of deer. It confines deer to relatively productive habitats, excluding them from deserts, dry grasslands, and geologically old landscapes leached of nutrients. Moreover, it severely limits the abundance of Cervidae in mature, species-rich faunas in which many herbivore species compete for food. In order to meet their high nutrient demands, deer are specialized to exploit disturbed ecosystems. For instance, after a forest fire, an area normally passes through several ecological plant successions within a few decades before the original conditions are restored. Early plant successions normally contain an abundance of the type of plant food required by deer. Some disturbances, such as river flooding and the rise and fall of lake levels, occur annually and create local, perpetually immature, nutrient-rich ecosystems. Since disturbances such as wildfires, storm floods, avalanches, or wind-felled trees are unpredictable, deer have evolved great abilities to quickly find and colonize such transient habitats. For example, the severe ecological upheaval caused by the extreme climatic oscillations of the Ice Ages greatly favoured deer. Glaciers ground rock into highly fertile waterborne silt and wind-borne loess that refertilized landscapes and rejuvenated the soil. Extinctions swept away warm-climate competitors. From the tropics deer spread to colder and more seasonal landscapes, including the Alps and the Arctic. Like other families of large mammals that colonized extreme Ice Age environments, deer diversified and evolved into grotesque giants that had ornate coat patterns and large, bizarre antlers, which could grow only from nutrient-rich soils.

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While deer tend to have broad, somewhat similar food habits, they are highly divergent in their antipredator strategies. This divergence segregates species ecologically and thus minimizes potential food competition between species sharing the same space. A deer species that hides and, if discovered, departs in rapid jumps to hide again requires forests and thickets, while a highly specialized runner needs flat, unobstructed terrain to outrun predators. Specialized jumpers may choose to stay close to steep slopes and rugged terrain and thus avoid areas frequented by species that run and jump, while cliff climbers may exploit gradients and altitudes closed to others.

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