ground beetle, (family Carabidae), any member of more than 40,000 insect species in one of the largest families in the insect order Coleoptera. They can be found in almost any terrestrial habitat on Earth. Ground beetles are recognized by their long legs and shiny black or brown elytra (wing covers), which are decorated with ridges and may be fused together along the midline. In many species the hind wings are reduced or absent. Ground beetles prefer moist cool areas and usually run rather than fly when disturbed. They emerge from under rocks, crevices, or litter at night in search of insects, worms, or snails. The long, slender larvae are mostly carnivorous, although those of a few species feed on seeds. They have sharp projecting mouthparts and a pair of bristly tail appendages. Many ground beetles secrete a foul-smelling liquid that discourages potential predators such as birds.

The searcher, or caterpillar hunter (Calosoma scrutator), is a common, brightly coloured North American ground beetle about 35 mm (1.5 inches) long. Its green or violet wings are edged in red, and its body has violet-blue, gold, and green markings. This and related species of ground beetles are known to climb trees in search of caterpillars. They secrete an acidic fluid that can blister human skin. The iridescent green-coloured C. sycophanta was introduced into North America from Europe to help control the spongy moth and brown-tail moth caterpillars.

The snail hunters (e.g., Scaphinotus) are a specialized group of ground beetles. Elongated, hook-shaped mouthparts allow them to extract the snail from its shell. The bombardier beetle (Brachinus in North America and Pherosophus in Africa, Asia, and the East Indies) has little sacs at the tip of its abdomen that spray a noxious fluid used to deter enemies. The beetle fires this fluid at boiling-hot temperatures, and the hot fluid vaporizes upon contact with air. The fluid itself consists of toxins called quinones that react explosively in an internal chamber with hydrogen peroxide, which is also produced by the beetle and stored in a separate body compartment. The expulsion of fluid from the abdomen is accompanied by a distinctive popping sound that may further startle the beetle’s predators. Brachinus species have dark blue, black, or blue-green wings and reddish yellow bodies and legs.

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The beneficial Lebia grandis, which resembles the bombardier beetle, preys upon the Colorado potato beetle. The Malayan leaf beetle, or fiddle beetle (Mormolyce), measuring approximately 100 mm (4 inches) long, resembles a violin with its slender head and thorax and wide elytra. This flat beetle uses its long head to probe into small openings in search of prey. It hides in crevices, under tree bark, and in porous fungi. The majority of ground beetles feed on insects harmful to humans and are therefore considered to be beneficial.

The Editors of Encyclopaedia Britannica This article was most recently revised and updated by Melissa Petruzzello.
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coleopteran, (order Coleoptera), any member of the insect order Coleoptera, consisting of the beetles and weevils. It is the largest order of insects, representing about 40 percent of the known insect species. Among the over 360,000 species of Coleoptera are many of the largest and most conspicuous insects, some of which also have brilliant metallic colours, showy patterns, or striking form. Beetles can usually be recognized by their two pairs of wings; the front pair is modified into horny covers (elytra) that hide the rear pair and most of the abdomen and usually meet down the back in a straight line. Coleoptera occur in nearly all climates. They may be divided into four groups: the first three, the Archostemata, the Adephaga, and the Myxophaga, contain relatively few families; the majority of beetles are placed in the fourth group, the Polyphaga.

Beetles attract attention for many different reasons, including their economic importance, size, abundance, appearance, and remarkable habits. Several groups of beetles (e.g., Lampyridae) are among the few terrestrial animals capable of producing light; members of several other families (e.g., Cerambycidae) can produce sound (stridulate). Most large beetles make a loud noise during flight, and many species, both large and small, are attracted to light at night. Some beetles (e.g., burying beetles of the family Silphidae and whirligig beetles of the family Gyrinidae) attract attention by their bizarre habits; others do so by their grotesque forms (e.g., Scarabaeidae). Many beetles have become adapted to an aquatic environment (e.g., Hydrophilidae); others (e.g., Thorictinae) live in association with ants and termites.

General features

Distribution and abundance

Beetles are found in nearly all climates and latitudes, except in such extreme environments as those in Antarctica and at the highest altitudes. They are found on subantarctic islands, close to the northern extremes in the Arctic, and on many mountaintops. Although many species occur in temperate environments, the number of species is greatest in the tropics; in general, individuals of a species are most abundant in temperate areas, and fewer individuals of more species are found in the tropics.

Some species are solitary; others occur in aggregations. Predators such as the ground beetles (Carabidae) are more apt to be found alone, as are many long-horned wood-boring beetles (Cerambycidae) and weevils (Curculionidae). Ladybugs (Coccinellidae), leaf beetles (Chrysomelidae), pleasing fungus beetles (Erotylidae), darkling beetles (Tenebrionidae), checkered beetles (Cleridae), bess beetles (Passalidae), sap beetles (Nitidulidae), and some species of scarab beetles (Scarabaeoidea) are often found in aggregations of one or several different species.

Most families contain both widely distributed species and some with very limited ranges. Wide distribution in this sense refers to a zoogeographical or faunal region; limited distribution, to a single valley, plain, island, altitude zone, or vegetation type on a mountain.

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Size range and diversity of structure

Coleoptera vary greatly in size, from a fraction of a millimetre to more than 200 mm (almost 8 inches) in length (e.g., rhinoceros beetle, Xyloryctes satyrus) and up to 75 mm (2.95 inches) in width (e.g., goliath beetle, Goliathus goliathus).

Diversity of structure among adult beetles is as great as range of size. The ground beetles (Carabidae) have a rather generalized (primitive) form—the flattened, oval body has a relatively even surface, with regular ridges or grooves; antennae and legs are of moderate length and slender. The underside of most water beetles (Hydrophilidae) is oval, smooth, and flattened, the antennae either short or very slender, and the forelegs short and the hindlegs long and fringed with hairs that are used as paddles. Rove beetles (Staphylinidae) have very short elytra and a slender abdomen. Soldier beetles (Cantharidae), fireflies (Lampyridae), and net-winged beetles (Lycidae) have soft elytra.

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Click beetles (Elateridae) have a hingelike joint in the body region called the thorax that enables them to snap their bodies and jump high in the air; their relatives, the Buprestidae (metallic wood borers), cannot jump but take flight very quickly. Cleridae (checkered beetles) are usually oblong or cylindrical, fairly active, and often brightly coloured. Nitidulidae (sap beetles) are short and flattened and have slightly shortened elytra. Coccinellidae (ladybugs, ladybird beetles) are rounded, with a smooth, raised upper surface and a flat underside. The Endomychidae (handsome fungus beetles) often have enlarged, rounded elytra. The Erotylidae (pleasing fungus beetles) are usually slender, smooth, and shiny, as are the Languriidae.

Among the stout or cylindrical lamellicorns (Scarabaeoidea) are a number of bizarre forms. The male rhinoceros beetles (Dynastinae) have one or more horns on the head and sometimes on part of the thorax. Many of the true scarabs (Scarabaeinae) and other dung-feeding groups of the lamellicorns also have horns, including some of the goliath beetles (Cetoniinae). Male stag beetles (Lucanidae) have greatly enlarged mandibles (jaws); some are as long as the rest of the body.

The Chrysomelidae (leaf beetles) vary from simple egg-shaped forms to slender, flat, or wedge-shaped ones, with wide elytra in the tortoise beetles and often numerous spines or tubercles in the leaf-mining leaf beetles (Cassidinae). The Bruchinae (seed beetles, or bean weevils) are short and stumpy, with short stout legs. The head, slightly elongated in front, is similar to that of some curculionid weevils. The Cerambycidae (long-horned beetles), diverse in form and structure, usually have antennae that are longer than the body. Cerambycids may be slender and medium to large in size or very small.

The extremely diverse Tenebrionidae (darkling beetles) are not always recognized as members of one family. Most arboreal (tree-dwelling) forms in the tropics or subtropics are slender and long-legged. Some slender and egg-shaped forms have a metallic sheen; most of the ground-dwelling forms are black and robust. The large tropical Trictenotomidae resemble some cerambycids (Prioninae) but are not related to them. The Alleculinae (comb-clawed beetles) resemble some of the slender tenebrionids but are usually more active. Lagriinae (lagriids) have a characteristic shape, usually widened behind, and sometimes a metallic sheen. Colydiinae (cylindrical bark beetles), hard-bodied and shiny or roughened, may be cylindrical and somewhat flattened.

The Curculionidae (weevils) range from slender to stout, elongated to egg-shaped; the bodies of some species contain many rounded projections (tubercles), and those of others may be smooth or grooved. The mouth is located on the end of a snoutlike projection, which varies in shape from short and stout to long and slender and sometimes exceeds the length of the rest of the body. Some Anthribidae (fungus weevils), usually cylindrical in shape, have slender antennae that may be longer than the rest of the body; they are easily confused with the cerambycids. Brentidae (primitive weevils) usually are long and slender with antennae projecting from the sides of the snout. Scolytinae (bark beetles, ambrosia beetles) do not have a distinct snout and are usually cylindrical in shape, as are Platypodinae.

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