langur, general name given to numerous species of Asian monkeys belonging to the subfamily Colobinae. The term is often restricted to nearly two dozen species of leaf monkeys but is also applied to various other members of the subfamily.

Leaf monkeys and other langurs are gregarious, diurnal, and basically arboreal monkeys with long tails and slender bodies. The limbs, hands, and feet are also long and slender. Depending on species, the head and body are about 40 to 80 cm (16 to 31 inches) long and the tail about 50 to 110 cm; weight varies from 5.5 kg (12 pounds) in the smallest species, the white-fronted langur (Presbytis frontata) of Borneo, up to 15 kg in the female and 19 kg in the male of the Himalayan langur (Semnopithecus schistaceus). Leaf monkeys have long fur, and many species have characteristic caps or crests of long hair. Colour varies among species but is commonly gray, red, brown, or black, and adults usually have black faces. The colour of the young, born singly after five to six months’ gestation, differs from that of adults and possibly serves to arouse the protective instincts of the adults. Mothers are protective but allow other females to help care for the young. Like the related colobus monkeys of Africa, langurs have large, complex stomachs adapted to a diet of leaves, fruit, and other vegetation.

The gray, or Hanuman, langur (S. entellus) of the Indian subcontinent is almost black when newborn and gray, tan, or brown as an adult. Regarded as sacred in Hinduism, it spends a good deal of time on the ground and roams at will in villages and temples of India and Nepal, raiding crops and the stores of merchants. The Hanuman langur usually lives in bands of about 20 to 30, though some troops number over 100. In some regions troops include several dominance-ranked adult males, though elsewhere there is only a single adult male per troop. In single-male troops, surplus males live in small bachelor bands that occasionally attempt to oust a troop leader. If successful, one of the bachelors takes over the troop and attempts to kill the unweaned infants in order to bring the females quickly back into estrus (mating condition).

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Leaf monkeys of genus Trachypithecus are also called brow-ridged langurs. They live in Southeast Asia from Bhutan and southern China to Java and are smaller and more arboreal than Hanuman langurs. The newborn are a bright golden colour. There are 10 to 15 species, including the beautiful golden langur (T. geei) from Bhutan, the spectacled langur (T. obscurus) from the Malay Peninsula, with white eye rings and pink muzzle, and a group of black langurs with white markings on the head and body, including François’ langur (T. francoisi) and its relatives, which live in the limestone country of northern Vietnam, Laos, and parts of southeastern China (Kwangsi). The purple-faced langur (T. vetulus) of Sri Lanka and the rare Nilgiri langur (T. johnii) of southern India may be more closely related to the Hanuman.

Leaf monkeys of the genus Presbytis are confined to Malaysia and western Indonesia, where they are mostly known by the local name surili; brow-ridged langurs of this region are generally called lutung. Most of the 10 or so Presbytis species are white on the underside and on the inner aspect of the thigh, contrasting sharply with the dark upper side. The newborn are white with a thick dark line from crown to rump and another at right angles across the shoulders (“cruciger” pattern). Most species live in small territorial groups of one male and two to four females with their young, but one, the joja (P. potenziani) of the Mentawai Islands of Indonesia, is unique among Old World monkeys in that it always lives in monogamous pairs.

The three genera of langurs commonly called leaf monkeys are Presbytis, Trachypithecus, and Semnopithecus; other langurs belong to the genera Pygathrix, Rhinopithecus, Nasalis, and Simias and include the proboscis monkey and simakobu. Several species are endangered. Langurs and colobus monkeys make up the subfamily Colobinae of the Old World monkey family, Cercopithecidae.

The Editors of Encyclopaedia Britannica This article was most recently revised and updated by John P. Rafferty.
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primate, in zoology, any mammal of the group that includes the lemurs, lorises, tarsiers, monkeys, apes, and humans. The order Primates, including more than 500 species, is the third most diverse order of mammals, after rodents (Rodentia) and bats (Chiroptera).

Although there are some notable variations between some primate groups, they share several anatomic and functional characteristics reflective of their common ancestry. When compared with body weight, the primate brain is larger than that of other terrestrial mammals, and it has a fissure unique to primates (the Calcarine sulcus) that separates the first and second visual areas on each side of the brain. Whereas all other mammals have claws or hooves on their digits, only primates have flat nails. Some primates do have claws, but even among these there is a flat nail on the big toe (hallux). In all primates except humans, the hallux diverges from the other toes and together with them forms a pincer capable of grasping objects such as branches. Not all primates have similarly dextrous hands; only the catarrhines (Old World monkeys, apes, and humans) and a few of the lemurs and lorises have an opposable thumb. Primates are not alone in having grasping feet, but as these occur in many other arboreal mammals (e.g., squirrels and opossums), and as most present-day primates are arboreal, this characteristic suggests that they evolved from an ancestor that was arboreal. So too does primates’ possession of specialized nerve endings (Meissner’s corpuscles) in the hands and feet that increase tactile sensitivity. As far as is known, no other placental mammal has them. Primates possess dermatoglyphics (the skin ridges responsible for fingerprints), but so do many other arboreal mammals.

The eyes face forward in all primates so that the eyes’ visual fields overlap. Again, this feature is not by any means restricted to primates, but it is a general feature seen among predators. It has been proposed, therefore, that the ancestor of the primates was a predator, perhaps insectivorous. The optic fibres in almost all mammals cross over (decussate) so that signals from one eye are interpreted on the opposite side of the brain, but, in some primate species, up to 40 percent of the nerve fibres do not cross over.

Primate teeth are distinguishable from those of other mammals by the low, rounded form of the molar and premolar cusps, which contrast with the high, pointed cusps or elaborate ridges of other placental mammals. This distinction makes fossilized primate teeth easy to recognize.

Fossils of the earliest primates date to the Early Eocene Epoch (56 million to 41.2 million years ago) or perhaps to the Late Paleocene Epoch (59.2 million to 56 million years ago). Though they began as an arboreal group, and many (especially the platyrrhines, or New World monkeys) have remained thoroughly arboreal, many have become at least partly terrestrial, and many have achieved high levels of intelligence. It is certainly no accident that the most intelligent of all forms of life, the only one capable of constructing the Encyclopædia Britannica, belongs to this order.

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By the 21st century the populations of approximately 75 percent of all primate species were falling, and some 60 percent were considered either threatened or endangered species. Habitat loss and fragmentation from logging, mining, urban sprawl, and the conversion of natural areas to agriculture and livestock raising are the primary threats to many species. Other causes of widespread population declines include hunting and poaching, the pet trade, the illegal trade in primate body parts, and the susceptibility of some primates to infection with human diseases.

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