Notoungulata, extinct group of hoofed mammals found as fossils, mostly in South America, although the oldest forms seem to have originated in East Asia. Notoungulates lived from the late Paleocene Epoch (about 57 million years ago) to the early part of the Pleistocene Epoch (some 1.8 million years ago) and were most diverse during the Miocene Epoch (23–5.3 million years ago). By the Pliocene Epoch (5.3–2.6 million years ago) their numbers and diversity were reduced, perhaps owing to changes in climate and geography. In South America, notoungulates evolved and diversified in isolation; they became extinct after the land connection between North and South America was reestablished about 3.5 million years ago.

In their time, the notoungulates included a variety of hoofed animals whose development paralleled the evolution of more-advanced forms elsewhere. One group, the toxodonts, was clumsily built and rather massive; Toxodon stood about 1.5 metres (5 feet) high at the shoulder. Other notoungulates developed along lines similar to rabbits and rodents.

This article was most recently revised and updated by Richard Pallardy.
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Paleogene Period

geochronology
Also known as: Palaeogene Period

Paleogene Period, oldest of the three stratigraphic divisions of the Cenozoic Era spanning the interval between 66 million and 23 million years ago. Paleogene is Greek meaning “ancient-born” and includes the Paleocene (Palaeocene) Epoch (66 million to 56 million years ago), the Eocene Epoch (56 million to 33.9 million years ago), and the Oligocene Epoch (33.9 million to 23 million years ago). The term Paleogene was devised in Europe to emphasize the similarity of marine fossils found in rocks of the first three Cenozoic epochs, as opposed to the later fossils of the Neogene Period (23 million to 2.6 million years ago) and the Quaternary Period (2.6 million years ago to the present). In North America, the Cenozoic has traditionally been divided only into the Tertiary Period (66 million to 2.6 million years ago) and the Quaternary Period; however, the notion that the Tertiary should be replaced by the designations Paleogene and Neogene is becoming more widespread.

This article was most recently revised and updated by John P. Rafferty.
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