oreodont, any member of a diverse group of extinct herbivorous North American artiodactyls (even-toed ungulates) that lived from the Middle Eocene through the end of the Miocene (between about 40 million and 5.3 million years ago). Though the best-known species, such as Leptauchenia and Merycoidodon, are often compared to sheep in size and shape, oreodonts are usually considered members of the suborder Typlopoda (the group that contains camels); however, some studies place them outside that group. Oreodonts were unlike any living mammal group in the structure of their skeleton and dentition. They diversified during the period when Earth’s climate was cooling from the Paleocene-Eocene Thermal Maximum (PETM) about 55.8 million years ago and reached their maximum diversity during the relatively cool Oligocene Epoch (34 million to 23 million years ago).

The earliest oreodonts belonged to the family Agriochoeridae. The diversity of these browsing, forest-dwelling mammals peaked in the late Eocene (between about 40 million and 34 million years ago). The most advanced agriochoerid, Agriochoerus, lived during the Oligocene, however. Later oreodonts, members of family Merycoidodontidae, had higher-crowned teeth that were specialized for tougher diets than the agriochoerids. The merycoidodonts were especially diverse; more than 19 genera are known, and 10 genera lived contemporaneously during the Miocene Epoch (23 million to 5 million years ago).

The skeletons of oreodonts were unusual compared with living artiodactyls in that they were not unguligrade (that is, walking habitually on their toes). Rather, oreodont skeletons supported a digitigrade stance (that is, their limbs were more like those of dogs and cats). In addition, the middle ears of some Oligocene oreodonts were also unusual in having extremely large chambers that appear to have been specialized for hearing low-frequency sounds.

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Oreodont fossils are especially common in the Brule Formation of the White River Badlands of South Dakota, U.S. This formation is composed of river deposits and paleosols (soils buried under sedimentary rock) that developed in savanna-like environments about 34 million years ago.

Paul David Polly
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fossil record, history of life as documented by fossils, the remains or imprints of organisms from earlier geological periods preserved in sedimentary rock. In a few cases the original substance of the hard parts of the organism is preserved, but more often the original components have been replaced by minerals deposited from water seeping through the rock. Occasionally the original material is simply removed while nothing is deposited in its place; in this case, all that remains is a mould of the shape of the plant or animal. A brief treatment of the fossil record follows. For full treatment, see geochronology.

In some places, such as the Grand Canyon in Arizona, it is possible to recognize a great thickness of nearly horizontal strata representing the deposition of sediment on the seafloor over many hundreds of millions of years. It is often observed that each layer in such a sequence contains fossils that are distinct from those of the layers that are above and below it. In such sequences of layers in different places, the same, or similar, fossil floras or faunas occur in the identical order. By comparison of overlapping sequences, it is possible to build up a continuous record of faunas or floras that have progressively more in common with present-day life-forms as the top of the sequence is approached.

Study of the fossil record has provided important information for at least three different purposes. The progressive changes observed within an animal group are used to describe the evolution of that group. In general, but not always, successive generations tend to change morphologically in a particular direction (e.g., the progressive acquisition or loss of specific features), and these changes are often interpreted as better adaptation (through preferential selection of beneficial mutations) to a particular environment.

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Fossils also provide the geologist a quick and easy way of assigning an age to the strata in which they occur. The precision with which this may be done in any particular case depends on the nature and abundance of the fauna: some fossil groups were deposited during much longer time intervals than others.

Fossil organisms, furthermore, may provide useful information about the climate and environment of the site where they were deposited and preserved. Certain species of coral, for example, require warm shallow water; certain plants require warm swampy conditions such as are found today in the Florida Everglades, South America’s Pantanal, and Botswana’s Okavango delta. Thus, when rocks containing fossils of this kind are found in rocks of the present-day polar regions, there is a strong presumption that the crust on which they were deposited has shifted its position on the surface of Earth since that time.

The Editors of Encyclopaedia BritannicaThis article was most recently revised and updated by Meg Matthias.
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