anther, in flowering plants, the part of a stamen that produces and contains pollen. Each anther is generally borne at the tip of a long slender stalk known as a filament and consists of two lobes that each house a pair of pollen sacs (microsporangia) that produce pollen for pollination. As the anther matures, the partition between the adjacent microsporangia of a pair breaks down so that there are only two pollen-containing sacs (one in each lobe of the anther) at the time the anther releases the pollen. While the anthers of most angiosperms dehisce, or release, pollen through a rupture along one side of each sac, those belonging to members of the heath family (Ericaceae) characteristically release pollen through small pores at the anther tip.