metacarpal, any of several tubular bones between the wrist (carpal) bones and each of the forelimb digits in land vertebrates, corresponding to the metatarsal bones of the foot. Originally numbering five, metacarpals in many mammals have undergone much change and reduction during evolution. The lower leg of the horse, for example, includes only one strengthened metacarpal; the two splint bones behind and above the hoof are reduced metacarpals, and the remaining two original metacarpals have been lost. In humans the five metacarpals are flat at the back of the hand and bowed on the palmar side; they form a longitudinal arch that accommodates the muscles, tendons, and nerves of the palm. The metacarpals also form a transverse arch that allows the fingertips and thumb to be brought together for manipulation.