thumb, short, thick first digit of the human hand and of the lower-primate hand and foot. It differs from other digits in having only two phalanges (tubular bones of the fingers and toes). The thumb also differs in having much freedom of movement and being opposable to tips of other digits. The corresponding first digit (most medial digit) in other vertebrates is also called the thumb, especially if it has some manipulative or special ability (e.g., in the raccoon and the bat).