plesiosaur, (clade Plesiosauria), any of a group of long-necked marine reptiles found as fossils from the late Triassic Period into the late Cretaceous Period (215 million to 66 million years ago). Plesiosaurs had a wide distribution in European seas and around the Pacific Ocean, including Australia, North America, and Asia.

Plesiosaurus, an early plesiosaur, was about 4.5 metres (15 feet) long, with a broad, flat body and a relatively short tail. It swam by flapping its fins in the water, much as sea lions do today, in a modified style of underwater “flight.” The nostrils were located far back on the head near the eyes. The neck was long and flexible, and the animal may have fed by swinging its head from side to side through schools of fish, capturing prey by using the long sharp teeth present in the jaws.

Early in their evolutionary history, the plesiosaurs split into two main lineages: the pliosaurs (or pliosauroids, which belong to the suborder Pliosauroidea), in which the neck was short and the head elongated; and the plesiosauroids (which belong to the suborder Plesiosauroidea), in which the head remained relatively small and the neck assumed snakelike proportions and became very flexible. The late evolution of plesiosaurs was marked by a great increase in size. For example, Elasmosaurus, a plesiosaurid, had as many as 76 vertebrae in its neck alone and reached a length of about 13 metres (43 feet), fully half of which consisted of the head and neck. In contrast, Kronosaurus, an early Cretaceous pliosaur whose fossils have been unearthed in Australia and Colombia, grew to about 9–11 metres (29.5–36 feet) long, based on an estimate of the length of K. queenlandicus (known by some researchers as Eiectus longmani) and K. boyacensis (known by some researchers as Monquirasaurus boyacensis); however, the reptile’s fossil skull alone measured between 2.2 and 2.9 metres (7.2 and 9.5 feet) long. An even larger pliosaur from the Jurassic Period, Pliosaurus funkei (known colloquially as “Predator X”), was unearthed in Svalbard in 2009. Its length and weight are estimated at 15 metres (about 50 feet) and 45 tonnes (almost 100,000 pounds), respectively. The jaws of this creature are thought to have produced a bite force of 33,000 psi (pound-force per square inch), perhaps the largest bite force of any known animal.

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fossil record, history of life as documented by fossils, the remains or imprints of organisms from earlier geological periods preserved in sedimentary rock. In a few cases the original substance of the hard parts of the organism is preserved, but more often the original components have been replaced by minerals deposited from water seeping through the rock. Occasionally the original material is simply removed while nothing is deposited in its place; in this case, all that remains is a mould of the shape of the plant or animal. A brief treatment of the fossil record follows. For full treatment, see geochronology.

In some places, such as the Grand Canyon in Arizona, it is possible to recognize a great thickness of nearly horizontal strata representing the deposition of sediment on the seafloor over many hundreds of millions of years. It is often observed that each layer in such a sequence contains fossils that are distinct from those of the layers that are above and below it. In such sequences of layers in different places, the same, or similar, fossil floras or faunas occur in the identical order. By comparison of overlapping sequences, it is possible to build up a continuous record of faunas or floras that have progressively more in common with present-day life-forms as the top of the sequence is approached.

Study of the fossil record has provided important information for at least three different purposes. The progressive changes observed within an animal group are used to describe the evolution of that group. In general, but not always, successive generations tend to change morphologically in a particular direction (e.g., the progressive acquisition or loss of specific features), and these changes are often interpreted as better adaptation (through preferential selection of beneficial mutations) to a particular environment.

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Fossils also provide the geologist a quick and easy way of assigning an age to the strata in which they occur. The precision with which this may be done in any particular case depends on the nature and abundance of the fauna: some fossil groups were deposited during much longer time intervals than others.

Fossil organisms, furthermore, may provide useful information about the climate and environment of the site where they were deposited and preserved. Certain species of coral, for example, require warm shallow water; certain plants require warm swampy conditions such as are found today in the Florida Everglades, South America’s Pantanal, and Botswana’s Okavango delta. Thus, when rocks containing fossils of this kind are found in rocks of the present-day polar regions, there is a strong presumption that the crust on which they were deposited has shifted its position on the surface of Earth since that time.

The Editors of Encyclopaedia BritannicaThis article was most recently revised and updated by Meg Matthias.
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