pliosaur, a group of large carnivorous marine reptiles characterized by massive heads, short necks, and streamlined tear-shaped bodies. Pliosaurs have been found as fossils from the Jurassic and Cretaceous periods (about 200 million to 65.5 million years ago). They are classified in the order Plesiosauria, along with their long-necked relatives, the plesiosaurs. Pliosaurs possessed powerful jaws and large teeth, and they used four large fins to swim through Mesozoic seas.

One notable pliosaur is Liopleurodon, a genus found in Middle Jurassic deposits in England and northern France. Liopleurodon is significant in that several fossils of variable quality that range in length from 5 to 25 metres (16 to 85 feet) have been placed in this genus, leading many authorities to question whether such specimens should be reclassified into other genera.

On the other hand, some groups did indeed grow quite large. For example, Kronosaurus, an Early Cretaceous pliosaur from Australia, grew to about 12 metres (about 40 feet) long; the skull alone measured about 3.7 metres (12.1 feet) long. An even larger pliosaur from the Jurassic, dubbed “Predator X,” was unearthed in Svalbard in 2009. Although it remains unclassified at present, some details are known. Its length and weight are estimated at 15 metres (about 50 feet) long and 45 tonnes (almost 100,000 pounds), respectively. The jaws of this creature are thought to have produced a bite force of 33,000 pounds per square inch, perhaps the highest bite force of any known animal.

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Another specimen, known from a massive skull unearthed from the coast of southern England, may be the longest pliosaur on record. Extrapolations made from the 2.4-metre (7.8-foot) skull suggest that the specimen ranged from 10 to 16 metres (33 to about 53 feet) from head to tail.

John P. Rafferty
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paleontology, scientific study of life of the geologic past that involves the analysis of plant and animal fossils, including those of microscopic size, preserved in rocks. It is concerned with all aspects of the biology of ancient life forms: their shape and structure, evolutionary patterns, taxonomic relationships with each other and with modern living species, geographic distribution, and interrelationships with the environment. Paleontology is mutually interdependent with stratigraphy and historical geology because fossils constitute a major means by which sedimentary strata are identified and correlated with one another. Its methods of investigation include that of biometry (statistical analysis applied to biology), which is designed to provide a description of the forms of organisms statistically and the expression of taxonomic relationships quantitatively.

Paleontology has played a key role in reconstructing Earth’s history and has provided much evidence to support the theory of evolution. Data from paleontological studies, moreover, have aided petroleum geologists in locating deposits of oil and natural gas. The occurrence of such fossil fuels is frequently associated with the presence of the remains of certain ancient life-forms.

Paleontological research dates back to the early 1800s. In 1815 the English geologist William Smith demonstrated the value of using fossils for the study of strata. About the same time, the French zoologist Georges Cuvier initiated comparative studies of the structure of living animals with fossil remains.

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The Editors of Encyclopaedia BritannicaThis article was most recently revised and updated by Adam Augustyn.
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