Also called:
large carpenter bee

carpenter bee, (genus Xylocopa), any of a genus of about 400 species of bees that are found in most areas of the world. Like most bees, carpenter bees do not produce honey, and they do not sting unless provoked. Although bees in Xylocopa are often considered pests because of their tunneling in structural wood such as that of buildings and fences, they are also important native pollinators that coevolved with the flowering plants in the areas in which they live. This makes them well adapted to the climates, flower structures, and other ecological relationships in their habitats, and it makes them essential to the perpetuation of many wild species.

Taxonomy

See also list of ants, bees, and wasps.

Physical description

Carpenter bees somewhat resemble bumblebees (Bombus) but differ in having a nonhairy abdomen. Most are all black or nearly so and may feature white or yellow hairs on the thorax. Males sometimes have a white or yellow face and often have larger compound eyes than females. Only the females have a stinger, which is a modified ovipositor for egg-laying.

Natural history

Most carpenter bees are solitary, meaning that each female builds and provisions her own brood nest. The bees are so named for their habit of nesting in a tunnel excavated within solid wood. A few species are ground nesting. Under certain environmental conditions, some species form simple social colonies of mothers and daughters and divide the labor of foraging and egg-laying among themselves.

Sea otter (Enhydra lutris), also called great sea otter, rare, completely marine otter of the northern Pacific, usually found in kelp beds. Floats on back. Looks like sea otter laughing. saltwater otters
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Commonly built into tree branches, wooden beams, or even park benches, carpenter bee nests each have a single entrance, often a circular hole, that typically branches into a number of adjacent tunnels. Although the female uses her strong mandibles to carve a new nest or to adapt an existing nest, the wood is not eaten. Once the nest is established, the female lays her eggs in individual cells in the tunnels. The eggs, which are among the largest insect eggs known, are each provisioned with food before the cell is sealed with a mixture of wood pulp and saliva. The larvae take several weeks to reach maturity and pupate within their cells. Adults typically hibernate over the winter in the nest in which they matured and emerge the following spring to mate and produce their own offspring. Carpenter bee larvae and adults are an important food for woodpeckers.

Unrelated species

Another group of bees, commonly called small carpenter bees, form the genus Ceratina (tribe Ceratinini). Adults are typically about 6 mm (0.2 inch) long and metallic in coloration. Most are solitary and nest in plant stems, which the female first hollows out and then packs with pollen and eggs. A number of individual cells are placed in a row, separated by thin partitions of wood debris mixed with saliva.

The Editors of Encyclopaedia Britannica Melissa Petruzzello
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pollination, transfer of pollen grains from the stamens (the flower parts that produce them) to the ovule-bearing organs or to the ovules (seed precursors) themselves. In gymnosperm plants such as conifers and cycads, in which the ovules are exposed, the pollen is simply caught in a drop of fluid secreted by the ovule. In flowering plants, however, the ovules are contained within a hollow organ called the pistil, and the pollen is deposited on the pistil’s receptive surface, the stigma. There the pollen germinates and gives rise to a pollen tube, which grows down through the pistil toward one of the ovules in its base. In an act of double fertilization, one of the two sperm cells within the pollen tube fuses with the egg cell of the ovule, making possible the development of an embryo, and the other cell combines with the two subsidiary sexual nuclei of the ovule, which initiates formation of a reserve food tissue, the endosperm. The growing ovule then transforms itself into a seed.

As a prerequisite for fertilization, pollination is essential to the perpetuation of the vast majority of the world’s wild plants as well as to the production of most fruit and seed crops. It also plays an important part in programs designed to improve plants by breeding. Furthermore, studies of pollination are invaluable for understanding the evolution of flowering plants and their distribution in the world today. As sedentary organisms, plants usually must enlist the services of external agents for pollen transport. In flowering plants, these are (roughly in order of diminishing importance) insects, wind, birds, mammals, and water. See also major types of pollinators.

Types: self-pollination and cross-pollination

An egg cell in an ovule of a flower may be fertilized by a sperm cell derived from a pollen grain produced by that same flower or by another flower on the same plant, in either of which two cases fertilization is said to be due to self-pollination (autogamy); or, the sperm may be derived from pollen originating on a different plant individual, in which case the process is called cross-pollination (heterogamy). Both processes are common, but cross-pollination clearly has certain evolutionary advantages for the species: the seeds formed may combine the hereditary traits of both parents, and the resulting offspring generally are more varied than would be the case after self-pollination. In a changing environment, some of the individuals resulting from cross-pollination still may be found capable of coping with their new situation, ensuring survival of the species, whereas the individuals resulting from self-pollination might all be unable to adjust. Self-pollination, or selfing, although foolproof in a stable environment, thus is an evolutionary cul-de-sac. There also is a more direct, visible difference between selfing and outbreeding (cross-pollination): in those species where both methods work, cross-pollination usually produces more, and better quality, seeds. A dramatic demonstration of this effect is found with hybrid corn (maize), a superior product that results from cross-breeding of several especially bred lines.

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